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Taking place between the events of the Game Boy Advance titles Mega Man Battle Network 1 and 2, this action platformer spin-off of the Mega Man Battle Network RPG series sees Lan and his virtual avatar Mega Man on a mission to stop an evil virus codenamed "Zero" from destroying the cyberspace. Gameplay involves guiding Mega Man through each side-scrolling level, hacking into cyberspace and battling infected victims of the virus. Players can choose from a range of weapons and special abilities stored on Battle Chips, which can be selected and downloaded once found. Each chip provides a new weapon or combat option for Mega Man. In addition to the usual boss battles at the end of certain levels and puzzle elements throughout, Network Transmission also features light RPG elements taken from the other Battle Network games where Mega Man's various attributes can be upgraded.
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